How did deadly Syria clashes start and who is responsible for civilian killings?

More than 1,000 people are believed to have died so far in fighting between security services and Assad loyalists
Clashes between Syrian security services and fighters loyal to the ousted Assad regime erupted on Wednesday, kicking off five days of still-ongoing fighting which has killed more than 1,000 people, including 745 civilians, according to a war monitor.
The clashes, some of the deadliest in the country since the beginning of its civil war in 2011, were the biggest challenge Syria’s new authorities faced since taking power in December.
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Hundreds of civilians killed in Syria as clashes erupt in former Assad strongholds
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Hundreds of civilians have been killed in Syria. Here's what we know.
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The UK-based Syrian Observatory for Human Rights (SOHR) said 830 civilians had been killed.BBC News - 1d -
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What We Know About the Clashes in Syria
Scores have been killed in fierce clashes between government forces and gunmen loyal to the Assad regime, in a serious challenge to the country’s new rulers.The New York Times - 2d
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